Borneo

At www.sciencedaily.com/releases/2019/04/190425073625.htm … a research paper in Scientific Reports (2019:9 (I) 10.1038/s41598-019-42670-4) claims there was a savannah grassland landscape in Borneo during the Late Pleistocene (presumably coinciding with the Late Glacial Maximum). Borneo is an equatorial forest in the modern world and has probably been that way for most of the Holocene. Savannah landscapes are usually located north or south of the tropical forest zone – in Africa for example. Not sure what the situation is in SE or southern Asia but could this be further evidence that in the LGM period the equator was in a different position to what it is today.